These are the best muffins!!!! I have used all different berries, and they come out great! My favorite I'd say are when I use fresh strawberries. I use 3/4 of a cup of each flour and substitute 1/4 cup of flax seed and 1/4 cup of wheat germ. Lately I've been making them vegan. Instead of egg I used a mashed banana. Instead of buttermilk I used almond milk and lemon juice. I use coconut oil instead of canola oil. Be sure to use an organic lemon since you are using the rind. I used organic cane sugar, its still sugar, but its not as porcessed or bleached, and to cut down on the sugar I halfed it, and used dark wildflower honey for the other half. I do sprinkle the top with just a little raw turbinado sugar though. Really these are fantastic!